// This model generated automatically from SBML

// unit definitions
import nsrunit;
unit conversion off;

// SBML property definitions
property sbmlRole=string;
property sbmlName=string;
property sbmlCompartment=string;

// SBML reactions
// vPTS: 65cpep cglcex => 65cg6p 65cpyr 
// vPGI: cg6p => cf6p 
// vPGM: cg6p => cg1p 
// vG6PDH: cg6p => cpg 
// vPFK: cf6p => cfdp 
// vTA: csed7p cgap => cf6p ce4p 
// vTKA: cxyl5p crib5p => cgap csed7p 
// vTKB: cxyl5p ce4p => cf6p cgap 
// vMURSyNTH: 2cf6p => X 
// vALDO: cfdp => cgap cdhap 
// vGAPDH: cgap => cpgp 
// vTIS: cdhap => cgap 
// vTRPSYNTH: X => cgap cpyr 
// vG3PDH: cdhap => X 
// vPGK: cpgp => cpg3 
// vsersynth: cpg3 
// vrpGluMu: cpg3 => cpg2 
// vENO: cpg2 => cpep 
// vPK: cpep => cpyr 
// vpepCxylase: cpep 
// vSynth1: cpep 
// vSynth2: cpyr 
// vDAHPS: ce4p cpep 
// vPDH: cpyr 
// vMethSynth: => cpyr 
// vPGDH: cpg 
// vR5PI: cribu5p => crib5p 
// vRu5P: cribu5p => cxyl5p 
// vPPK: crib5p => X 
// vG1PAT: cg1p => X 
// vG6P: cg6p => X 
// vf6P: cf6p => X 
// vfdP: cfdp => X 
// vGAP: cgap => X 
// vDHAP: cdhap => X 
// vPGP: cpgp => X 
// vPG3: cpg3 => X 
// vpg2: cpg2 => X 
// vPEP: cpep => X 
// vRibu5p: cribu5p => X 
// vRIB5P: crib5p => X 
// vXYL5P: cxyl5p => X 
// vSED7P: csed7p => X 
// vpyr: cpyr => X 
// vPG: cpg => X 
// vE4P: ce4p => X 
// vGLP: cg1p => X 
// vEXTER: X => cglcex 

math main {
  realDomain time second;
  time.min=0;
  extern time.max;
  extern time.delta;

  // variable definitions
  real compartment = 1 L;
  real rmaxPTS = 7829.78;
  real rmaxTKa = 9.4733848;
  real rmaxENO = 330.4476151;
  real rmaxSynth1 = .01953897;
  real rmaxPGluMu = 89.04965407;
  real rmaxPK = .0611315;
  real rmaxPDH = 6.059531;
  real rmaxG1PAT = .00752546;
  real rmaxSerSynth = .02571211;
  real rmaxDAHPS = .10795312;
  real rmaxPGK = 3021.773771;
  real rmaxRu5P = 6.7390295;
  real rmaxR5PI = 4.8384119;
  real rmaxPGM = .83982428;
  real rmaxRPPK = .01290045;
  real rmaxG6PDH = 1.380197;
  real rmaxALDO = 17.41464425;
  real rmaxTIS = 68.67474392;
  real rmaxG3PDH = .01162043;
  real rmaxPFK = 1840.584747;
  real rmaxPGI = 650.9878687;
  real rmaxPGDH = 1.623236E1;
  real rmaxGAPDH = 921.5942861;
  real rmaxSynth2 = .07361855;
  real rmaxTKb = 86.55855855;
  real rmaxTA = 10.87164108;
  real rmaxpepCxylase = .10702059;
  real rmaxMurSynth = 4.3711E-4;
  real rmaxTrpSynth = .001037;
  real rmaxMetSynth = .0022627;
  real Dil = 2.78E-5;
  real mu = 2.78E-5;
  real cfeed = 110.96;
  real KTKbeq = 10;
  real KPGIg6p = 2.9;
  real KPGIeq = .1725;
  real KPGIf6p = .266;
  real KPGIg6ppginh = .2;
  real KPGIf6ppginh = .2;
  real KSynth1pep = 1;
  real KGAPDHeq = .63;
  real KGAPDHgap = .683;
  real KGAPDHnad = .252;
  real KGAPDHpgp = 1.04E-5;
  real KGAPDHnadh = 1.09;
  real KRu5Peq = 1.4;
  real KG1PATg1p = 3.2;
  real KG1PATfdp = .119;
  real KG1PATatp = 4.42;
  real nG1PATfdp = 1.2;
  real KR5PIeq = 4;
  real KPGKeq = 1934.4;
  real KPGKadp = .185;
  real KPGKpgp = .0468;
  real KPGKatp = .653;
  real KPGKpg3 = .473;
  real KENOeq = 6.73;
  real KENOpg2 = .1;
  real KENOpep = .135;
  real nPK = 4;
  real LPK = 1E3;
  real KTAeq = 1.05;
  real KG6PDHnadp = .0246;
  real KG6PDHnadphnadpinh = .01;
  real KG6PDHnadphg6pinh = 6.43;
  real KG6PDHg6p = 14.4;
  real KPTSa1 = 3082.3;
  real KPTSa2 = .01;
  real KPTSa3 = 245.3;
  real KPTSg6p = 2.15;
  real nPTSg6p = 3.66;
  real KPKpep = .31;
  real KPKatp = 22.5;
  real KPKadp = .26;
  real KPKamp = .2;
  real KPKfdp = .19;
  real KSerSynthpg3 = 1;
  real KPDHpyr = 1159;
  real nPDH = 3.68;
  real KPGluMueq = .188;
  real KPGluMupg3 = .2;
  real KPGluMupg2 = .369;
  real kTISeq = 1.39;
  real kTISdhap = 2.8;
  real kTISgap = .3;
  real KPGDHpg = 37.5;
  real KPGDHnadp = .0506;
  real KPGDHatpinh = 208;
  real KPGDHnadphinh = .0138;
  real KPGMeq = .196;
  real KPGMg6p = 1.038;
  real KPGMg1p = .0136;
  real KRPPKrib5p = .1;
  real KpepCxylasepep = 4.07;
  real KpepCxylasefdp = .7;
  real npepCxylasefdp = 4.21;
  real KG3PDHdhap = 1;
  real KPFKf6ps = .325;
  real KPFKatps = .123;
  real KPFKpep = 3.26;
  real KPFKadpa = 128;
  real KPFKadpc = 4.14;
  real KPFKadpb = 3.89;
  real KPFKampa = 19.1;
  real KPFKampb = 3.2;
  real nPFK = 11.1;
  real KTKaeq = 1.2;
  real KDAHPSe4p = .035;
  real KDAHPSpep = .0053;
  real nDAHPSe4p = 2.6;
  real nDAHPSpep = 2.2;
  real LPFK = 5629067;
  real kALDOgap = .088;
  real kALDOgapinh = .6;
  real kALDOdhap = .088;
  real VALDOblf = 2;
  real KSynth2pyr = 1;
  real kALDOeq = .144;
  real kALDOfdp = 1.75;
  real cnad = 1.47;
  real cnadh = .1;
  real catp = 4.27;
  real cadp = .595;
  real camp = .955;
  real cnadp = .195;
  real cnadph = .062;
  real cpep(time) M;
  real cglcex(time) M;
  real cg6p(time) M;
  real cpyr(time) M;
  real cf6p(time) M;
  real cpg(time) M;
  real cg1p(time) M;
  real cfdp(time) M;
  real csed7p(time) M;
  real cgap(time) M;
  real ce4p(time) M;
  real cxyl5p(time) M;
  real crib5p(time) M;
  real X = 1 M;
  real cdhap(time) M;
  real cpgp(time) M;
  real cpg3(time) M;
  real cpg2(time) M;
  real cribu5p(time) M;
  real vPTS(time) katal;
  real vPGI(time) katal;
  real vPGM(time) katal;
  real vG6PDH(time) katal;
  real vPFK(time) katal;
  real vTA(time) katal;
  real vTKA(time) katal;
  real vTKB(time) katal;
  real vMURSyNTH(time) katal;
  real vALDO(time) katal;
  real vGAPDH(time) katal;
  real vTIS(time) katal;
  real vTRPSYNTH(time) katal;
  real vG3PDH(time) katal;
  real vPGK(time) katal;
  real vsersynth(time) katal;
  real vrpGluMu(time) katal;
  real vENO(time) katal;
  real vPK(time) katal;
  real vpepCxylase(time) katal;
  real vSynth1(time) katal;
  real vSynth2(time) katal;
  real vDAHPS(time) katal;
  real vPDH(time) katal;
  real vMethSynth(time) katal;
  real vPGDH(time) katal;
  real vR5PI(time) katal;
  real vRu5P(time) katal;
  real vPPK(time) katal;
  real vG1PAT(time) katal;
  real vG6P(time) katal;
  real vf6P(time) katal;
  real vfdP(time) katal;
  real vGAP(time) katal;
  real vDHAP(time) katal;
  real vPGP(time) katal;
  real vPG3(time) katal;
  real vpg2(time) katal;
  real vPEP(time) katal;
  real vRibu5p(time) katal;
  real vRIB5P(time) katal;
  real vXYL5P(time) katal;
  real vSED7P(time) katal;
  real vpyr(time) katal;
  real vPG(time) katal;
  real vE4P(time) katal;
  real vGLP(time) katal;
  real vEXTER(time) katal;

  // equations
  when (time=time.min) cpep = 2.67;
  cpep:time = (-65*vPTS + vENO + -1*vPK + -1*vpepCxylase + -1*vSynth1 + -1*vDAHPS + -1*vPEP)/compartment;
  when (time=time.min) cglcex = .0556;
  cglcex:time = (-1*vPTS + vEXTER)/compartment;
  when (time=time.min) cg6p = 3.48;
  cg6p:time = (65*vPTS + -1*vPGI + -1*vPGM + -1*vG6PDH + -1*vG6P)/compartment;
  when (time=time.min) cpyr = 2.67;
  cpyr:time = (65*vPTS + vTRPSYNTH + vPK + -1*vSynth2 + -1*vPDH + vMethSynth + -1*vpyr)/compartment;
  when (time=time.min) cf6p = .6;
  cf6p:time = (vPGI + -1*vPFK + vTA + vTKB + -2*vMURSyNTH + -1*vf6P)/compartment;
  when (time=time.min) cpg = .8075;
  cpg:time = (vG6PDH + -1*vPGDH + -1*vPG)/compartment;
  when (time=time.min) cg1p = .6525;
  cg1p:time = (vPGM + -1*vG1PAT + -1*vGLP)/compartment;
  when (time=time.min) cfdp = .272;
  cfdp:time = (vPFK + -1*vALDO + -1*vfdP)/compartment;
  when (time=time.min) csed7p = .276;
  csed7p:time = (-1*vTA + vTKA + -1*vSED7P)/compartment;
  when (time=time.min) cgap = .218;
  cgap:time = (-1*vTA + vTKA + vTKB + vALDO + -1*vGAPDH + vTIS + vTRPSYNTH + -1*vGAP)/compartment;
  when (time=time.min) ce4p = .098;
  ce4p:time = (vTA + -1*vTKB + -1*vDAHPS + -1*vE4P)/compartment;
  when (time=time.min) cxyl5p = .138;
  cxyl5p:time = (-1*vTKA + -1*vTKB + vRu5P + -1*vXYL5P)/compartment;
  when (time=time.min) crib5p = .398;
  crib5p:time = (-1*vTKA + vR5PI + -1*vPPK + -1*vRIB5P)/compartment;
  when (time=time.min) cdhap = .167;
  cdhap:time = (vALDO + -1*vTIS + -1*vG3PDH + -1*vDHAP)/compartment;
  when (time=time.min) cpgp = .008;
  cpgp:time = (vGAPDH + -1*vPGK + -1*vPGP)/compartment;
  when (time=time.min) cpg3 = 2.131;
  cpg3:time = (vPGK + -1*vsersynth + -1*vrpGluMu + -1*vPG3)/compartment;
  when (time=time.min) cpg2 = .399;
  cpg2:time = (vrpGluMu + -1*vENO + -1*vpg2)/compartment;
  when (time=time.min) cribu5p = .111;
  cribu5p:time = (-1*vR5PI + -1*vRu5P + -1*vRibu5p)/compartment;
  vPTS = rmaxPTS*cglcex*cpep/cpyr/(KPTSa1+KPTSa2*cpep/cpyr+KPTSa3*cglcex+cglcex*cpep/cpyr)/(1+cg6p^nPTSg6p/KPTSg6p);
  vPGI = rmaxPGI*(cg6p-cf6p/KPGIeq)/(KPGIg6p*(1+cf6p/KPGIf6p/(1+cpg/KPGIf6ppginh)+cpg/KPGIg6ppginh)+cg6p);
  vPGM = rmaxPGM*(cg6p-cg1p/KPGMeq)/(KPGMg6p*(1+cg1p/KPGMg1p)+cg6p);
  vG6PDH = rmaxG6PDH*cg6p*cnadp/(cg6p+KG6PDHg6p)/(1+cnadph/KG6PDHnadphg6pinh)/(KG6PDHnadp*(1+cnadph/KG6PDHnadphnadpinh)+cnadp);
  vPFK = rmaxPFK*catp/(catp+KPFKatps*(1+cadp/KPFKadpc))*cf6p/(cf6p+KPFKf6ps*(1+cpep/KPFKpep+cadp/KPFKadpb+camp/KPFKampb)/(1+cadp/KPFKadpa+camp/KPFKampa))/(1+LPFK/(1+cf6p/KPFKf6ps/(1+cpep/KPFKpep+cadp/KPFKadpb+camp/KPFKampb)*(1+cadp/KPFKadpa+camp/KPFKampa))^nPFK);
  vTA = rmaxTA*(cgap*csed7p-ce4p*cf6p/KTAeq);
  vTKA = rmaxTKa*(crib5p*cxyl5p-csed7p*cgap/KTKaeq);
  vTKB = rmaxTKb*(cxyl5p*ce4p-cf6p*cgap/KTKbeq);
  vMURSyNTH = rmaxMurSynth;
  vALDO = rmaxALDO*(cfdp-cgap*cdhap/kALDOeq)/(kALDOfdp+cfdp+kALDOgap/kALDOeq/VALDOblf*cdhap+kALDOdhap/kALDOeq/VALDOblf*cgap+cfdp*cgap/kALDOgapinh+cgap*cdhap/VALDOblf/kALDOeq);
  vGAPDH = rmaxGAPDH*(cgap*cnad-cpgp*cnadh/KGAPDHeq)/(KGAPDHgap*(1+cpgp/KGAPDHpgp)+cgap)/(KGAPDHnad*(1+cnadh/KGAPDHnadh)+cnad);
  vTIS = rmaxTIS*(cdhap-cgap/kTISeq)/(kTISdhap*(1+cgap/kTISgap)+cdhap);
  vTRPSYNTH = rmaxTrpSynth;
  vG3PDH = rmaxG3PDH*cdhap/(KG3PDHdhap+cdhap);
  vPGK = rmaxPGK*(cadp*cpgp-catp*cpg3/KPGKeq)/(KPGKadp*(1+catp/KPGKatp)+cadp)/(KPGKpgp*(1+cpg3/KPGKpg3)+cpgp);
  vsersynth = rmaxSerSynth*cpg3/(KSerSynthpg3+cpg3);
  vrpGluMu = rmaxPGluMu*(cpg3-cpg2/KPGluMueq)/(KPGluMupg3*(1+cpg2/KPGluMupg2)+cpg3);
  vENO = rmaxENO*(cpg2-cpep/KENOeq)/(KENOpg2*(1+cpep/KENOpep)+cpg2);
  vPK = rmaxPK*cpep/KPKpep*(cpep/KPKpep+1)^(nPK-1)/(LPK*((1+catp/KPKatp)/(cfdp/KPKfdp+camp/KPKamp+1))^nPK+(cpep/KPKpep+1)^nPK)*cadp/(cadp+KPKadp);
  vpepCxylase = rmaxpepCxylase*cpep*(1+(cfdp/KpepCxylasefdp)^npepCxylasefdp)/(KpepCxylasepep+cpep);
  vSynth1 = rmaxSynth1*cpep/(KSynth1pep+cpep);
  vSynth2 = rmaxSynth2*cpyr/(KSynth2pyr+cpyr);
  vDAHPS = rmaxDAHPS*ce4p^nDAHPSe4p*cpep^nDAHPSpep/(KDAHPSe4p+ce4p^nDAHPSe4p)/(KDAHPSpep+cpep^nDAHPSpep);
  vPDH = rmaxPDH*cpyr^nPDH/(KPDHpyr+cpyr^nPDH);
  vMethSynth = rmaxMetSynth;
  vPGDH = rmaxPGDH*cpg*cnadp/(cpg+KPGDHpg)/(cnadp+KPGDHnadp*(1+cnadph/KPGDHnadphinh)*(1+catp/KPGDHatpinh));
  vR5PI = rmaxR5PI*(cribu5p-crib5p/KR5PIeq);
  vRu5P = rmaxRu5P*(cribu5p-cxyl5p/KRu5Peq);
  vPPK = rmaxRPPK*crib5p/(KRPPKrib5p+crib5p);
  vG1PAT = rmaxG1PAT*cg1p*catp*(1+(cfdp/KG1PATfdp)^nG1PATfdp)/(KG1PATatp+catp)/(KG1PATg1p+cg1p);
  vG6P = mu*cg6p;
  vf6P = mu*cf6p;
  vfdP = mu*cfdp;
  vGAP = mu*cgap;
  vDHAP = mu*cdhap;
  vPGP = mu*cpgp;
  vPG3 = mu*cpg3;
  vpg2 = mu*cpg2;
  vPEP = mu*cpep;
  vRibu5p = mu*cribu5p;
  vRIB5P = mu*crib5p;
  vXYL5P = mu*cxyl5p;
  vSED7P = mu*csed7p;
  vpyr = mu*cpyr;
  vPG = mu*cpg;
  vE4P = mu*ce4p;
  vGLP = mu*cg1p;
  vEXTER = Dil*(cfeed-cglcex);

  // variable properties
  compartment.sbmlRole="compartment";
  rmaxPTS.sbmlRole="parameter";
  rmaxTKa.sbmlRole="parameter";
  rmaxENO.sbmlRole="parameter";
  rmaxSynth1.sbmlRole="parameter";
  rmaxPGluMu.sbmlRole="parameter";
  rmaxPK.sbmlRole="parameter";
  rmaxPDH.sbmlRole="parameter";
  rmaxG1PAT.sbmlRole="parameter";
  rmaxSerSynth.sbmlRole="parameter";
  rmaxDAHPS.sbmlRole="parameter";
  rmaxPGK.sbmlRole="parameter";
  rmaxRu5P.sbmlRole="parameter";
  rmaxR5PI.sbmlRole="parameter";
  rmaxPGM.sbmlRole="parameter";
  rmaxRPPK.sbmlRole="parameter";
  rmaxG6PDH.sbmlRole="parameter";
  rmaxALDO.sbmlRole="parameter";
  rmaxTIS.sbmlRole="parameter";
  rmaxG3PDH.sbmlRole="parameter";
  rmaxPFK.sbmlRole="parameter";
  rmaxPGI.sbmlRole="parameter";
  rmaxPGDH.sbmlRole="parameter";
  rmaxGAPDH.sbmlRole="parameter";
  rmaxSynth2.sbmlRole="parameter";
  rmaxTKb.sbmlRole="parameter";
  rmaxTA.sbmlRole="parameter";
  rmaxpepCxylase.sbmlRole="parameter";
  rmaxMurSynth.sbmlRole="parameter";
  rmaxTrpSynth.sbmlRole="parameter";
  rmaxMetSynth.sbmlRole="parameter";
  Dil.sbmlRole="parameter";
  mu.sbmlRole="parameter";
  cfeed.sbmlRole="parameter";
  KTKbeq.sbmlRole="parameter";
  KPGIg6p.sbmlRole="parameter";
  KPGIeq.sbmlRole="parameter";
  KPGIf6p.sbmlRole="parameter";
  KPGIg6ppginh.sbmlRole="parameter";
  KPGIf6ppginh.sbmlRole="parameter";
  KSynth1pep.sbmlRole="parameter";
  KGAPDHeq.sbmlRole="parameter";
  KGAPDHgap.sbmlRole="parameter";
  KGAPDHnad.sbmlRole="parameter";
  KGAPDHpgp.sbmlRole="parameter";
  KGAPDHnadh.sbmlRole="parameter";
  KRu5Peq.sbmlRole="parameter";
  KG1PATg1p.sbmlRole="parameter";
  KG1PATfdp.sbmlRole="parameter";
  KG1PATatp.sbmlRole="parameter";
  nG1PATfdp.sbmlRole="parameter";
  KR5PIeq.sbmlRole="parameter";
  KPGKeq.sbmlRole="parameter";
  KPGKadp.sbmlRole="parameter";
  KPGKpgp.sbmlRole="parameter";
  KPGKatp.sbmlRole="parameter";
  KPGKpg3.sbmlRole="parameter";
  KENOeq.sbmlRole="parameter";
  KENOpg2.sbmlRole="parameter";
  KENOpep.sbmlRole="parameter";
  nPK.sbmlRole="parameter";
  LPK.sbmlRole="parameter";
  KTAeq.sbmlRole="parameter";
  KG6PDHnadp.sbmlRole="parameter";
  KG6PDHnadphnadpinh.sbmlRole="parameter";
  KG6PDHnadphg6pinh.sbmlRole="parameter";
  KG6PDHg6p.sbmlRole="parameter";
  KPTSa1.sbmlRole="parameter";
  KPTSa2.sbmlRole="parameter";
  KPTSa3.sbmlRole="parameter";
  KPTSg6p.sbmlRole="parameter";
  nPTSg6p.sbmlRole="parameter";
  KPKpep.sbmlRole="parameter";
  KPKatp.sbmlRole="parameter";
  KPKadp.sbmlRole="parameter";
  KPKamp.sbmlRole="parameter";
  KPKfdp.sbmlRole="parameter";
  KSerSynthpg3.sbmlRole="parameter";
  KPDHpyr.sbmlRole="parameter";
  nPDH.sbmlRole="parameter";
  KPGluMueq.sbmlRole="parameter";
  KPGluMupg3.sbmlRole="parameter";
  KPGluMupg2.sbmlRole="parameter";
  kTISeq.sbmlRole="parameter";
  kTISdhap.sbmlRole="parameter";
  kTISgap.sbmlRole="parameter";
  KPGDHpg.sbmlRole="parameter";
  KPGDHnadp.sbmlRole="parameter";
  KPGDHatpinh.sbmlRole="parameter";
  KPGDHnadphinh.sbmlRole="parameter";
  KPGMeq.sbmlRole="parameter";
  KPGMg6p.sbmlRole="parameter";
  KPGMg1p.sbmlRole="parameter";
  KRPPKrib5p.sbmlRole="parameter";
  KpepCxylasepep.sbmlRole="parameter";
  KpepCxylasefdp.sbmlRole="parameter";
  npepCxylasefdp.sbmlRole="parameter";
  KG3PDHdhap.sbmlRole="parameter";
  KPFKf6ps.sbmlRole="parameter";
  KPFKatps.sbmlRole="parameter";
  KPFKpep.sbmlRole="parameter";
  KPFKadpa.sbmlRole="parameter";
  KPFKadpc.sbmlRole="parameter";
  KPFKadpb.sbmlRole="parameter";
  KPFKampa.sbmlRole="parameter";
  KPFKampb.sbmlRole="parameter";
  nPFK.sbmlRole="parameter";
  KTKaeq.sbmlRole="parameter";
  KDAHPSe4p.sbmlRole="parameter";
  KDAHPSpep.sbmlRole="parameter";
  nDAHPSe4p.sbmlRole="parameter";
  nDAHPSpep.sbmlRole="parameter";
  LPFK.sbmlRole="parameter";
  kALDOgap.sbmlRole="parameter";
  kALDOgapinh.sbmlRole="parameter";
  kALDOdhap.sbmlRole="parameter";
  VALDOblf.sbmlRole="parameter";
  KSynth2pyr.sbmlRole="parameter";
  kALDOeq.sbmlRole="parameter";
  kALDOfdp.sbmlRole="parameter";
  cnad.sbmlRole="parameter";
  cnadh.sbmlRole="parameter";
  catp.sbmlRole="parameter";
  cadp.sbmlRole="parameter";
  camp.sbmlRole="parameter";
  cnadp.sbmlRole="parameter";
  cnadph.sbmlRole="parameter";
  cpep.sbmlRole="species";
  cpep.sbmlCompartment="compartment";
  cglcex.sbmlRole="species";
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}