// This model generated automatically from SBML

// unit definitions
import nsrunit;
unit conversion off;

// SBML property definitions
property sbmlRole=string;
property sbmlName=string;
property sbmlCompartment=string;

// SBML reactions
// R05640: Lanosterol <=> space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol 
// R04427: Zymosterol <=> Fecosterol 
// R02874: Squalene <=> space_S_space_minus_2_space_3_minus_Epoxysqualene 
// R02872: Presqualene_space_diphosphate <=> Squalene 
// R03199: space_S_space_minus_2_space_3_minus_Epoxysqualene <=> Lanosterol 
// R07509: space_14_minus_Demethyllanosterol <=> space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate 
// R02003: Geranyl_space_diphosphate Isopentenyl_space_diphosphate <=> trans_space_trans_minus_Farnesyl_space_diphosphate 
// R07483: space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol <=> space_4_alpha_minus_Methylfecosterol 
// R05639: space_14_minus_Demethyllanosterol <=> space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol 
// R02082: space_R_space_minus_Mevalonate <=> space_S_space_minus_3_minus_Hydroxy_minus_3_minus_methylglutaryl_minus_CoA 
// R05731: Obtusifoliol <=> space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol 
// R01123: Isopentenyl_space_diphosphate => Dimethylallyl_space_diphosphate 
// R05641: Ergosterol <=> Ergosta_minus_5_space_7_space_22_space_24_space_28_space_minus_tetraen_minus_3_beta_minus_ol 
// R07494: space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate <=> space_3_minus_Keto_minus_4_minus_methylzymosterol 
// R02245: space_R_space_minus_Mevalonate <=> space_R_space_minus_5_minus_Phosphomevalonate 
// R03245: space_R_space_minus_5_minus_Phosphomevalonate <=> space_R_space_minus_5_minus_Diphosphomevalonate 
// R07481: Cycloartenol <=> space_24_minus_Methylenecycloartanol 
// R01121: space_R_space_minus_5_minus_Diphosphomevalonate <=> Isopentenyl_space_diphosphate 
// R01658: Dimethylallyl_space_diphosphate Isopentenyl_space_diphosphate <=> Geranyl_space_diphosphate 
// R07495: space_3_minus_Keto_minus_4_minus_methylzymosterol <=> space_4_alpha_minus_Methylzymosterol 
// R00702: trans_space_trans_minus_Farnesyl_space_diphosphate <=> Presqualene_space_diphosphate 

math main {
  realDomain time second;
  time.min=0;
  extern time.max;
  extern time.delta;

  // variable definitions
  real default = 1 L;
  real uVol = 1 L;
  real E5_space_3_space_3_space_2 = 0 M;
  real E2_space_5_space_1_space_10 = 0 M;
  real E4_space_1_space_1_space_33 = 0 M;
  real E2_space_5_space_1_space_21 = 0 M;
  real E1_space_14_space_99_space_7 = 0 M;
  real E5_space_4_space_99_space_7 = 0 M;
  real E1_space_14_space_13_space_70 = 0 M;
  real E1_space_3_space_1_space_70 = 0 M;
  real E1_space_14_space_13_space_72 = 0 M;
  real E1_space_1_space_1_space_170 = 0 M;
  real E1_space_1_space_1_space_270 = 0 M;
  real E2_space_1_space_1_space_41 = 0 M;
  real E2_space_7_space_4_space_2 = 0 M;
  real E1_space_3_space_1_space_71 = 0 M;
  real E2_space_7_space_1_space_36 = 0 M;
  real E1_space_1_space_1_space_34 = 0 M;
  real Dimethylallyl_space_diphosphate(time) M;
  real Geranyl_space_diphosphate(time) M;
  real Isopentenyl_space_diphosphate(time) M;
  real trans_space_trans_minus_Farnesyl_space_diphosphate(time) M;
  real Presqualene_space_diphosphate(time) M;
  real Squalene(time) M;
  real space_S_space_minus_2_space_3_minus_Epoxysqualene(time) M;
  real Lanosterol(time) M;
  real space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol(time) M;
  real space_14_minus_Demethyllanosterol(time) M;
  real space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate(time) M;
  real space_3_minus_Keto_minus_4_minus_methylzymosterol(time) M;
  real space_4_alpha_minus_Methylzymosterol(time) M;
  real Zymosterol(time) M;
  real space_R_space_minus_5_minus_Diphosphomevalonate(time) M;
  real space_R_space_minus_5_minus_Phosphomevalonate(time) M;
  real Ergosta_minus_5_space_7_space_22_space_24_space_28_space_minus_tetraen_minus_3_beta_minus_ol(time) M;
  real Ergosterol(time) M;
  real Fecosterol(time) M;
  real Cycloartenol(time) M;
  real space_R_space_minus_Mevalonate(time) M;
  real space_24_minus_Methylenecycloartanol(time) M;
  real Obtusifoliol(time) M;
  real space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol(time) M;
  real space_4_alpha_minus_Methylfecosterol(time) M;
  real space_S_space_minus_3_minus_Hydroxy_minus_3_minus_methylglutaryl_minus_CoA(time) M;
  real R05640(time) katal;
  extern real R05640.kf;
  extern real R05640.kb;
  real R04427(time) katal;
  extern real R04427.kf;
  extern real R04427.kb;
  real R02874(time) katal;
  extern real R02874.kf;
  extern real R02874.kb;
  real R02872(time) katal;
  extern real R02872.kf;
  extern real R02872.kb;
  real R03199(time) katal;
  extern real R03199.kf;
  extern real R03199.kb;
  real R07509(time) katal;
  extern real R07509.kf;
  extern real R07509.kb;
  real R02003(time) katal;
  extern real R02003.kf;
  extern real R02003.kb;
  real R07483(time) katal;
  extern real R07483.kf;
  extern real R07483.kb;
  real R05639(time) katal;
  extern real R05639.kf;
  extern real R05639.kb;
  real R02082(time) katal;
  extern real R02082.kf;
  extern real R02082.kb;
  real R05731(time) katal;
  extern real R05731.kf;
  extern real R05731.kb;
  real R01123(time) katal;
  extern real R01123.kf;
  extern real R01123.kb;
  real R05641(time) katal;
  extern real R05641.kf;
  extern real R05641.kb;
  real R07494(time) katal;
  extern real R07494.kf;
  extern real R07494.kb;
  real R02245(time) katal;
  extern real R02245.kf;
  extern real R02245.kb;
  real R03245(time) katal;
  extern real R03245.kf;
  extern real R03245.kb;
  real R07481(time) katal;
  extern real R07481.kf;
  extern real R07481.kb;
  real R01121(time) katal;
  extern real R01121.kf;
  extern real R01121.kb;
  real R01658(time) katal;
  extern real R01658.kf;
  extern real R01658.kb;
  real R07495(time) katal;
  extern real R07495.kf;
  extern real R07495.kb;
  real R00702(time) katal;
  extern real R00702.kf;
  extern real R00702.kb;

  // equations
  when (time=time.min) Dimethylallyl_space_diphosphate = 0;
  Dimethylallyl_space_diphosphate:time = (R01123 + -1*R01658)/uVol;
  when (time=time.min) Geranyl_space_diphosphate = 0;
  Geranyl_space_diphosphate:time = (-1*R02003 + R01658)/uVol;
  when (time=time.min) Isopentenyl_space_diphosphate = 0;
  Isopentenyl_space_diphosphate:time = (-1*R02003 + -1*R01123 + R01121 + -1*R01658)/uVol;
  when (time=time.min) trans_space_trans_minus_Farnesyl_space_diphosphate = 0;
  trans_space_trans_minus_Farnesyl_space_diphosphate:time = (R02003 + -1*R00702)/uVol;
  when (time=time.min) Presqualene_space_diphosphate = 0;
  Presqualene_space_diphosphate:time = (-1*R02872 + R00702)/uVol;
  when (time=time.min) Squalene = 0;
  Squalene:time = (-1*R02874 + R02872)/uVol;
  when (time=time.min) space_S_space_minus_2_space_3_minus_Epoxysqualene = 0;
  space_S_space_minus_2_space_3_minus_Epoxysqualene:time = (R02874 + -1*R03199)/uVol;
  when (time=time.min) Lanosterol = 0;
  Lanosterol:time = (-1*R05640 + R03199)/uVol;
  when (time=time.min) space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol = 0;
  space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol:time = (R05640 + R05639)/uVol;
  when (time=time.min) space_14_minus_Demethyllanosterol = 0;
  space_14_minus_Demethyllanosterol:time = (-1*R07509 + -1*R05639)/uVol;
  when (time=time.min) space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate = 0;
  space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate:time = (R07509 + -1*R07494)/uVol;
  when (time=time.min) space_3_minus_Keto_minus_4_minus_methylzymosterol = 0;
  space_3_minus_Keto_minus_4_minus_methylzymosterol:time = (R07494 + -1*R07495)/uVol;
  when (time=time.min) space_4_alpha_minus_Methylzymosterol = 0;
  space_4_alpha_minus_Methylzymosterol:time = (R07495)/uVol;
  when (time=time.min) Zymosterol = 0;
  Zymosterol:time = (-1*R04427)/uVol;
  when (time=time.min) space_R_space_minus_5_minus_Diphosphomevalonate = 0;
  space_R_space_minus_5_minus_Diphosphomevalonate:time = (R03245 + -1*R01121)/uVol;
  when (time=time.min) space_R_space_minus_5_minus_Phosphomevalonate = 0;
  space_R_space_minus_5_minus_Phosphomevalonate:time = (R02245 + -1*R03245)/uVol;
  when (time=time.min) Ergosta_minus_5_space_7_space_22_space_24_space_28_space_minus_tetraen_minus_3_beta_minus_ol = 0;
  Ergosta_minus_5_space_7_space_22_space_24_space_28_space_minus_tetraen_minus_3_beta_minus_ol:time = (R05641)/uVol;
  when (time=time.min) Ergosterol = 0;
  Ergosterol:time = (-1*R05641)/uVol;
  when (time=time.min) Fecosterol = 0;
  Fecosterol:time = (R04427)/uVol;
  when (time=time.min) Cycloartenol = 0;
  Cycloartenol:time = (-1*R07481)/uVol;
  when (time=time.min) space_R_space_minus_Mevalonate = 0;
  space_R_space_minus_Mevalonate:time = (-1*R02082 + -1*R02245)/uVol;
  when (time=time.min) space_24_minus_Methylenecycloartanol = 0;
  space_24_minus_Methylenecycloartanol:time = (R07481)/uVol;
  when (time=time.min) Obtusifoliol = 0;
  Obtusifoliol:time = (-1*R05731)/uVol;
  when (time=time.min) space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol = 0;
  space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol:time = (-1*R07483 + R05731)/uVol;
  when (time=time.min) space_4_alpha_minus_Methylfecosterol = 0;
  space_4_alpha_minus_Methylfecosterol:time = (R07483)/uVol;
  when (time=time.min) space_S_space_minus_3_minus_Hydroxy_minus_3_minus_methylglutaryl_minus_CoA = 0;
  space_S_space_minus_3_minus_Hydroxy_minus_3_minus_methylglutaryl_minus_CoA:time = (R02082)/uVol;
  R05640 = R05640.kf*Lanosterol - R05640.kb*space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol;
  R04427 = R04427.kf*Zymosterol - R04427.kb*Fecosterol;
  R02874 = R02874.kf*Squalene - R02874.kb*space_S_space_minus_2_space_3_minus_Epoxysqualene;
  R02872 = R02872.kf*Presqualene_space_diphosphate - R02872.kb*Squalene;
  R03199 = R03199.kf*space_S_space_minus_2_space_3_minus_Epoxysqualene - R03199.kb*Lanosterol;
  R07509 = R07509.kf*space_14_minus_Demethyllanosterol - R07509.kb*space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate;
  R02003 = R02003.kf*Geranyl_space_diphosphate*Isopentenyl_space_diphosphate - R02003.kb*trans_space_trans_minus_Farnesyl_space_diphosphate;
  R07483 = R07483.kf*space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol - R07483.kb*space_4_alpha_minus_Methylfecosterol;
  R05639 = R05639.kf*space_14_minus_Demethyllanosterol - R05639.kb*space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol;
  R02082 = R02082.kf*space_R_space_minus_Mevalonate - R02082.kb*space_S_space_minus_3_minus_Hydroxy_minus_3_minus_methylglutaryl_minus_CoA;
  R05731 = R05731.kf*Obtusifoliol - R05731.kb*space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol;
  R01123 = R01123.kf*Isopentenyl_space_diphosphate - R01123.kb*Dimethylallyl_space_diphosphate;
  R05641 = R05641.kf*Ergosterol - R05641.kb*Ergosta_minus_5_space_7_space_22_space_24_space_28_space_minus_tetraen_minus_3_beta_minus_ol;
  R07494 = R07494.kf*space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate - R07494.kb*space_3_minus_Keto_minus_4_minus_methylzymosterol;
  R02245 = R02245.kf*space_R_space_minus_Mevalonate - R02245.kb*space_R_space_minus_5_minus_Phosphomevalonate;
  R03245 = R03245.kf*space_R_space_minus_5_minus_Phosphomevalonate - R03245.kb*space_R_space_minus_5_minus_Diphosphomevalonate;
  R07481 = R07481.kf*Cycloartenol - R07481.kb*space_24_minus_Methylenecycloartanol;
  R01121 = R01121.kf*space_R_space_minus_5_minus_Diphosphomevalonate - R01121.kb*Isopentenyl_space_diphosphate;
  R01658 = R01658.kf*Dimethylallyl_space_diphosphate*Isopentenyl_space_diphosphate - R01658.kb*Geranyl_space_diphosphate;
  R07495 = R07495.kf*space_3_minus_Keto_minus_4_minus_methylzymosterol - R07495.kb*space_4_alpha_minus_Methylzymosterol;
  R00702 = R00702.kf*trans_space_trans_minus_Farnesyl_space_diphosphate - R00702.kb*Presqualene_space_diphosphate;

  // variable properties
  default.sbmlRole="compartment";
  uVol.sbmlRole="compartment";
  E5_space_3_space_3_space_2.sbmlRole="species";
  E5_space_3_space_3_space_2.sbmlCompartment="uVol";
  E2_space_5_space_1_space_10.sbmlRole="species";
  E2_space_5_space_1_space_10.sbmlCompartment="uVol";
  E4_space_1_space_1_space_33.sbmlRole="species";
  E4_space_1_space_1_space_33.sbmlCompartment="uVol";
  E2_space_5_space_1_space_21.sbmlRole="species";
  E2_space_5_space_1_space_21.sbmlCompartment="uVol";
  E1_space_14_space_99_space_7.sbmlRole="species";
  E1_space_14_space_99_space_7.sbmlCompartment="uVol";
  E5_space_4_space_99_space_7.sbmlRole="species";
  E5_space_4_space_99_space_7.sbmlCompartment="uVol";
  E1_space_14_space_13_space_70.sbmlRole="species";
  E1_space_14_space_13_space_70.sbmlCompartment="uVol";
  E1_space_3_space_1_space_70.sbmlRole="species";
  E1_space_3_space_1_space_70.sbmlCompartment="uVol";
  E1_space_14_space_13_space_72.sbmlRole="species";
  E1_space_14_space_13_space_72.sbmlCompartment="uVol";
  E1_space_1_space_1_space_170.sbmlRole="species";
  E1_space_1_space_1_space_170.sbmlCompartment="uVol";
  E1_space_1_space_1_space_270.sbmlRole="species";
  E1_space_1_space_1_space_270.sbmlCompartment="uVol";
  E2_space_1_space_1_space_41.sbmlRole="species";
  E2_space_1_space_1_space_41.sbmlCompartment="uVol";
  E2_space_7_space_4_space_2.sbmlRole="species";
  E2_space_7_space_4_space_2.sbmlCompartment="uVol";
  E1_space_3_space_1_space_71.sbmlRole="species";
  E1_space_3_space_1_space_71.sbmlCompartment="uVol";
  E2_space_7_space_1_space_36.sbmlRole="species";
  E2_space_7_space_1_space_36.sbmlCompartment="uVol";
  E1_space_1_space_1_space_34.sbmlRole="species";
  E1_space_1_space_1_space_34.sbmlCompartment="uVol";
  Dimethylallyl_space_diphosphate.sbmlRole="species";
  Dimethylallyl_space_diphosphate.sbmlCompartment="uVol";
  Geranyl_space_diphosphate.sbmlRole="species";
  Geranyl_space_diphosphate.sbmlCompartment="uVol";
  Isopentenyl_space_diphosphate.sbmlRole="species";
  Isopentenyl_space_diphosphate.sbmlCompartment="uVol";
  trans_space_trans_minus_Farnesyl_space_diphosphate.sbmlRole="species";
  trans_space_trans_minus_Farnesyl_space_diphosphate.sbmlCompartment="uVol";
  Presqualene_space_diphosphate.sbmlRole="species";
  Presqualene_space_diphosphate.sbmlCompartment="uVol";
  Squalene.sbmlRole="species";
  Squalene.sbmlCompartment="uVol";
  space_S_space_minus_2_space_3_minus_Epoxysqualene.sbmlRole="species";
  space_S_space_minus_2_space_3_minus_Epoxysqualene.sbmlCompartment="uVol";
  Lanosterol.sbmlRole="species";
  Lanosterol.sbmlCompartment="uVol";
  space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol.sbmlRole="species";
  space_4_space_4_minus_Dimethyl_minus_5_alpha_minus_cholesta_minus_8_space_14_space_24_minus_trien_minus_3_beta_minus_ol.sbmlCompartment="uVol";
  space_14_minus_Demethyllanosterol.sbmlRole="species";
  space_14_minus_Demethyllanosterol.sbmlCompartment="uVol";
  space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate.sbmlRole="species";
  space_4_alpha_minus_Methylzymosterol_minus_4_minus_carboxylate.sbmlCompartment="uVol";
  space_3_minus_Keto_minus_4_minus_methylzymosterol.sbmlRole="species";
  space_3_minus_Keto_minus_4_minus_methylzymosterol.sbmlCompartment="uVol";
  space_4_alpha_minus_Methylzymosterol.sbmlRole="species";
  space_4_alpha_minus_Methylzymosterol.sbmlCompartment="uVol";
  Zymosterol.sbmlRole="species";
  Zymosterol.sbmlCompartment="uVol";
  space_R_space_minus_5_minus_Diphosphomevalonate.sbmlRole="species";
  space_R_space_minus_5_minus_Diphosphomevalonate.sbmlCompartment="uVol";
  space_R_space_minus_5_minus_Phosphomevalonate.sbmlRole="species";
  space_R_space_minus_5_minus_Phosphomevalonate.sbmlCompartment="uVol";
  Ergosta_minus_5_space_7_space_22_space_24_space_28_space_minus_tetraen_minus_3_beta_minus_ol.sbmlRole="species";
  Ergosta_minus_5_space_7_space_22_space_24_space_28_space_minus_tetraen_minus_3_beta_minus_ol.sbmlCompartment="uVol";
  Ergosterol.sbmlRole="species";
  Ergosterol.sbmlCompartment="uVol";
  Fecosterol.sbmlRole="species";
  Fecosterol.sbmlCompartment="uVol";
  Cycloartenol.sbmlRole="species";
  Cycloartenol.sbmlCompartment="uVol";
  space_R_space_minus_Mevalonate.sbmlRole="species";
  space_R_space_minus_Mevalonate.sbmlCompartment="uVol";
  space_24_minus_Methylenecycloartanol.sbmlRole="species";
  space_24_minus_Methylenecycloartanol.sbmlCompartment="uVol";
  Obtusifoliol.sbmlRole="species";
  Obtusifoliol.sbmlCompartment="uVol";
  space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol.sbmlRole="species";
  space_4_alpha_minus_Methyl_minus_5_alpha_minus_ergosta_minus_8_space_14_space_24_space_28_space_minus_trien_minus_3_beta_minus_ol.sbmlCompartment="uVol";
  space_4_alpha_minus_Methylfecosterol.sbmlRole="species";
  space_4_alpha_minus_Methylfecosterol.sbmlCompartment="uVol";
  space_S_space_minus_3_minus_Hydroxy_minus_3_minus_methylglutaryl_minus_CoA.sbmlRole="species";
  space_S_space_minus_3_minus_Hydroxy_minus_3_minus_methylglutaryl_minus_CoA.sbmlCompartment="uVol";
  R05640.sbmlRole="rate";
  R05640.kf.sbmlRole="rate";
  R05640.kb.sbmlRole="rate";
  R04427.sbmlRole="rate";
  R04427.kf.sbmlRole="rate";
  R04427.kb.sbmlRole="rate";
  R02874.sbmlRole="rate";
  R02874.kf.sbmlRole="rate";
  R02874.kb.sbmlRole="rate";
  R02872.sbmlRole="rate";
  R02872.kf.sbmlRole="rate";
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