// This model generated automatically from SBML

// unit definitions
import nsrunit;
unit conversion off;

// SBML property definitions
property sbmlRole=string;
property sbmlName=string;
property sbmlCompartment=string;

// SBML reactions
// R04990: space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone <=> space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone 
// R04986: space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate <=> space_2_minus_Octaprenylphenol 
// R04984: space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone <=> space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone 
// R04993: space_2_minus_Demethylmenaquinone <=> Menaquinone 
// R04988: space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol <=> space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol 
// R04989: space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol <=> space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone 
// R06859: space_2_minus_Phytyl_minus_1_space_4_minus_naphthoquinone <=> Phylloquinone 
// R05614: space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone => Ubiquinone 
// R02166: Ubiquinone => Ubiquinol 
// R05615: all_minus_trans_minus_Octaprenyl_space_diphosphate space_4_minus_Hydroxybenzoate <=> space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate 
// R04987: space_2_minus_Octaprenylphenol <=> space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol 

math main {
  realDomain time second;
  time.min=0;
  extern time.max;
  extern time.delta;

  // variable definitions
  real default = 1 L;
  real uVol = 1 L;
  real E1_space_14_space_13_space_minus_ = 0 M;
  real E1_space_6_space_5_space_3 = 0 M;
  real E4_space_1_space_1_space_minus_ = 0 M;
  real E2_space_1_space_1_space_64 = 0 M;
  real E2_space_1_space_1_space_minus_ = 0 M;
  real E2_space_5_space_1_space_minus_ = 0 M;
  real space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone(time) M;
  real space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone(time) M;
  real space_4_minus_Hydroxybenzoate(time) M;
  real Ubiquinol(time) M;
  real Ubiquinone(time) M;
  real space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate(time) M;
  real space_2_minus_Octaprenylphenol(time) M;
  real space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol(time) M;
  real space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol(time) M;
  real space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone(time) M;
  real space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone(time) M;
  real space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone(time) M;
  real all_minus_trans_minus_Octaprenyl_space_diphosphate(time) M;
  real space_2_minus_Demethylmenaquinone(time) M;
  real Menaquinone(time) M;
  real space_2_minus_Phytyl_minus_1_space_4_minus_naphthoquinone(time) M;
  real Phylloquinone(time) M;
  real R04990(time) katal;
  extern real R04990.kf;
  extern real R04990.kb;
  real R04986(time) katal;
  extern real R04986.kf;
  extern real R04986.kb;
  real R04984(time) katal;
  extern real R04984.kf;
  extern real R04984.kb;
  real R04993(time) katal;
  extern real R04993.kf;
  extern real R04993.kb;
  real R04988(time) katal;
  extern real R04988.kf;
  extern real R04988.kb;
  real R04989(time) katal;
  extern real R04989.kf;
  extern real R04989.kb;
  real R06859(time) katal;
  extern real R06859.kf;
  extern real R06859.kb;
  real R05614(time) katal;
  extern real R05614.kf;
  extern real R05614.kb;
  real R02166(time) katal;
  extern real R02166.kf;
  extern real R02166.kb;
  real R05615(time) katal;
  extern real R05615.kf;
  extern real R05615.kb;
  real R04987(time) katal;
  extern real R04987.kf;
  extern real R04987.kb;

  // equations
  when (time=time.min) space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone = 0;
  space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone:time = (-1*R04984)/uVol;
  when (time=time.min) space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone = 0;
  space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone:time = (R04984)/uVol;
  when (time=time.min) space_4_minus_Hydroxybenzoate = 0;
  space_4_minus_Hydroxybenzoate:time = (-1*R05615)/uVol;
  when (time=time.min) Ubiquinol = 0;
  Ubiquinol:time = (R02166)/uVol;
  when (time=time.min) Ubiquinone = 0;
  Ubiquinone:time = (R05614 + -1*R02166)/uVol;
  when (time=time.min) space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate = 0;
  space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate:time = (-1*R04986 + R05615)/uVol;
  when (time=time.min) space_2_minus_Octaprenylphenol = 0;
  space_2_minus_Octaprenylphenol:time = (R04986 + -1*R04987)/uVol;
  when (time=time.min) space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol = 0;
  space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol:time = (-1*R04988 + R04987)/uVol;
  when (time=time.min) space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol = 0;
  space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol:time = (R04988 + -1*R04989)/uVol;
  when (time=time.min) space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone = 0;
  space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone:time = (-1*R04990 + R04989)/uVol;
  when (time=time.min) space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone = 0;
  space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone:time = (R04990)/uVol;
  when (time=time.min) space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone = 0;
  space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone:time = (-1*R05614)/uVol;
  when (time=time.min) all_minus_trans_minus_Octaprenyl_space_diphosphate = 0;
  all_minus_trans_minus_Octaprenyl_space_diphosphate:time = (-1*R05615)/uVol;
  when (time=time.min) space_2_minus_Demethylmenaquinone = 0;
  space_2_minus_Demethylmenaquinone:time = (-1*R04993)/uVol;
  when (time=time.min) Menaquinone = 0;
  Menaquinone:time = (R04993)/uVol;
  when (time=time.min) space_2_minus_Phytyl_minus_1_space_4_minus_naphthoquinone = 0;
  space_2_minus_Phytyl_minus_1_space_4_minus_naphthoquinone:time = (-1*R06859)/uVol;
  when (time=time.min) Phylloquinone = 0;
  Phylloquinone:time = (R06859)/uVol;
  R04990 = R04990.kf*space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone - R04990.kb*space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone;
  R04986 = R04986.kf*space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate - R04986.kb*space_2_minus_Octaprenylphenol;
  R04984 = R04984.kf*space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone - R04984.kb*space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone;
  R04993 = R04993.kf*space_2_minus_Demethylmenaquinone - R04993.kb*Menaquinone;
  R04988 = R04988.kf*space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol - R04988.kb*space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol;
  R04989 = R04989.kf*space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol - R04989.kb*space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone;
  R06859 = R06859.kf*space_2_minus_Phytyl_minus_1_space_4_minus_naphthoquinone - R06859.kb*Phylloquinone;
  R05614 = R05614.kf*space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone - R05614.kb*Ubiquinone;
  R02166 = R02166.kf*Ubiquinone - R02166.kb*Ubiquinol;
  R05615 = R05615.kf*all_minus_trans_minus_Octaprenyl_space_diphosphate*space_4_minus_Hydroxybenzoate - R05615.kb*space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate;
  R04987 = R04987.kf*space_2_minus_Octaprenylphenol - R04987.kb*space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol;

  // variable properties
  default.sbmlRole="compartment";
  uVol.sbmlRole="compartment";
  E1_space_14_space_13_space_minus_.sbmlRole="species";
  E1_space_14_space_13_space_minus_.sbmlCompartment="uVol";
  E1_space_6_space_5_space_3.sbmlRole="species";
  E1_space_6_space_5_space_3.sbmlCompartment="uVol";
  E4_space_1_space_1_space_minus_.sbmlRole="species";
  E4_space_1_space_1_space_minus_.sbmlCompartment="uVol";
  E2_space_1_space_1_space_64.sbmlRole="species";
  E2_space_1_space_1_space_64.sbmlCompartment="uVol";
  E2_space_1_space_1_space_minus_.sbmlRole="species";
  E2_space_1_space_1_space_minus_.sbmlCompartment="uVol";
  E2_space_5_space_1_space_minus_.sbmlRole="species";
  E2_space_5_space_1_space_minus_.sbmlCompartment="uVol";
  space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlRole="species";
  space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlCompartment="uVol";
  space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlRole="species";
  space_2_minus_Hexaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlCompartment="uVol";
  space_4_minus_Hydroxybenzoate.sbmlRole="species";
  space_4_minus_Hydroxybenzoate.sbmlCompartment="uVol";
  Ubiquinol.sbmlRole="species";
  Ubiquinol.sbmlCompartment="uVol";
  Ubiquinone.sbmlRole="species";
  Ubiquinone.sbmlCompartment="uVol";
  space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate.sbmlRole="species";
  space_3_minus_Octaprenyl_minus_4_minus_hydroxybenzoate.sbmlCompartment="uVol";
  space_2_minus_Octaprenylphenol.sbmlRole="species";
  space_2_minus_Octaprenylphenol.sbmlCompartment="uVol";
  space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol.sbmlRole="species";
  space_2_minus_Octaprenyl_minus_6_minus_hydroxyphenol.sbmlCompartment="uVol";
  space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol.sbmlRole="species";
  space_2_minus_Octaprenyl_minus_6_minus_methoxyphenol.sbmlCompartment="uVol";
  space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlRole="species";
  space_2_minus_Octaprenyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlCompartment="uVol";
  space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlRole="species";
  space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlCompartment="uVol";
  space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlRole="species";
  space_2_minus_Octaprenyl_minus_3_minus_methyl_minus_5_minus_hydroxy_minus_6_minus_methoxy_minus_1_space_4_minus_benzoquinone.sbmlCompartment="uVol";
  all_minus_trans_minus_Octaprenyl_space_diphosphate.sbmlRole="species";
  all_minus_trans_minus_Octaprenyl_space_diphosphate.sbmlCompartment="uVol";
  space_2_minus_Demethylmenaquinone.sbmlRole="species";
  space_2_minus_Demethylmenaquinone.sbmlCompartment="uVol";
  Menaquinone.sbmlRole="species";
  Menaquinone.sbmlCompartment="uVol";
  space_2_minus_Phytyl_minus_1_space_4_minus_naphthoquinone.sbmlRole="species";
  space_2_minus_Phytyl_minus_1_space_4_minus_naphthoquinone.sbmlCompartment="uVol";
  Phylloquinone.sbmlRole="species";
  Phylloquinone.sbmlCompartment="uVol";
  R04990.sbmlRole="rate";
  R04990.kf.sbmlRole="rate";
  R04990.kb.sbmlRole="rate";
  R04986.sbmlRole="rate";
  R04986.kf.sbmlRole="rate";
  R04986.kb.sbmlRole="rate";
  R04984.sbmlRole="rate";
  R04984.kf.sbmlRole="rate";
  R04984.kb.sbmlRole="rate";
  R04993.sbmlRole="rate";
  R04993.kf.sbmlRole="rate";
  R04993.kb.sbmlRole="rate";
  R04988.sbmlRole="rate";
  R04988.kf.sbmlRole="rate";
  R04988.kb.sbmlRole="rate";
  R04989.sbmlRole="rate";
  R04989.kf.sbmlRole="rate";
  R04989.kb.sbmlRole="rate";
  R06859.sbmlRole="rate";
  R06859.kf.sbmlRole="rate";
  R06859.kb.sbmlRole="rate";
  R05614.sbmlRole="rate";
  R05614.kf.sbmlRole="rate";
  R05614.kb.sbmlRole="rate";
  R02166.sbmlRole="rate";
  R02166.kf.sbmlRole="rate";
  R02166.kb.sbmlRole="rate";
  R05615.sbmlRole="rate";
  R05615.kf.sbmlRole="rate";
  R05615.kb.sbmlRole="rate";
  R04987.sbmlRole="rate";
  R04987.kf.sbmlRole="rate";
  R04987.kb.sbmlRole="rate";
}