// This model generated automatically from SBML

// unit definitions
import nsrunit;
unit conversion off;

// SBML property definitions
property sbmlRole=string;
property sbmlName=string;
property sbmlCompartment=string;

// SBML reactions
// R03652: L_minus_Glutamine <=> Glutaminyl_minus_tRNA 
// R00257: L_minus_Glutamine <=> NAD_plus_ 
// R00261: L_minus_Glutamate <=> space_4_minus_Aminobutanoate 
// R01648: space_4_minus_Aminobutanoate => Succinate_space_semialdehyde 
// R00894: L_minus_Glutamate <=> gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine 
// R01072: space_5_minus_Phosphoribosylamine <=> L_minus_Glutamine 
// R00248: L_minus_Glutamate => space_2_minus_Oxoglutarate 
// R00256: L_minus_Glutamine => L_minus_Glutamate NH3 
// R00575: L_minus_Glutamine <=> Carbamoyl_space_phosphate 
// R00497: gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine <=> Glutathione 
// R00768: L_minus_Glutamine <=> D_minus_Glucosamine_space_6_minus_phosphate 
// R05578: L_minus_Glutamate <=> L_minus_Glutamyl_minus_tRNA_space_Glu_space_ 
// R00355: space_2_minus_Oxoglutarate => L_minus_Glutamate 
// R00253: L_minus_Glutamate => L_minus_Glutamine 
// R00094: Glutathione => Glutathione_space_disulfide 
// R00713: Succinate_space_semialdehyde => Succinate 
// R01231: L_minus_Glutamine <=> GMP 
// R00243: L_minus_Glutamate => space_2_minus_Oxoglutarate 
// R00114: L_minus_Glutamate <=> L_minus_Glutamine space_2_minus_Oxoglutarate 

math main {
  realDomain time second;
  time.min=0;
  extern time.max;
  extern time.delta;

  // variable definitions
  real default = 1 L;
  real uVol = 1 L;
  real E1_space_8_space_1_space_7 = 0 M;
  real E2_space_6_space_1_space_19 = 0 M;
  real E6_space_3_space_2_space_3 = 0 M;
  real E1_space_4_space_1_space_13 = 0 M;
  real E1_space_4_space_1_space_3 = 0 M;
  real E1_space_4_space_1_space_4 = 0 M;
  real E2_space_6_space_1_space_1 = 0 M;
  real E6_space_3_space_2_space_2 = 0 M;
  real E6_space_1_space_1_space_17 = 0 M;
  real E4_space_1_space_1_space_15 = 0 M;
  real E6_space_3_space_1_space_2 = 0 M;
  real E4_space_1_space_1_space_19 = 0 M;
  real E6_space_3_space_5_space_1 = 0 M;
  real E6_space_3_space_5_space_2 = 0 M;
  real E2_space_6_space_1_space_16 = 0 M;
  real E2_space_4_space_2_space_14 = 0 M;
  real E3_space_5_space_1_space_2 = 0 M;
  real E6_space_3_space_5_space_5 = 0 M;
  real E6_space_1_space_1_space_18 = 0 M;
  real E1_space_2_space_1_space_16 = 0 M;
  real Succinate(time) M;
  real Succinate_space_semialdehyde(time) M;
  real space_2_minus_Oxoglutarate(time) M;
  real Glutathione_space_disulfide(time) M;
  real Glutathione(time) M;
  real gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine(time) M;
  real L_minus_Glutamyl_minus_tRNA_space_Glu_space_(time) M;
  real space_4_minus_Aminobutanoate(time) M;
  real L_minus_Glutamine(time) M;
  real L_minus_Glutamate(time) M;
  real space_5_minus_Phosphoribosylamine(time) M;
  real NAD_plus_(time) M;
  real GMP(time) M;
  real D_minus_Glucosamine_space_6_minus_phosphate(time) M;
  real Carbamoyl_space_phosphate(time) M;
  real NH3(time) M;
  real Glutaminyl_minus_tRNA(time) M;
  real R03652(time) katal;
  extern real R03652.kf;
  extern real R03652.kb;
  real R00257(time) katal;
  extern real R00257.kf;
  extern real R00257.kb;
  real R00261(time) katal;
  extern real R00261.kf;
  extern real R00261.kb;
  real R01648(time) katal;
  extern real R01648.kf;
  extern real R01648.kb;
  real R00894(time) katal;
  extern real R00894.kf;
  extern real R00894.kb;
  real R01072(time) katal;
  extern real R01072.kf;
  extern real R01072.kb;
  real R00248(time) katal;
  extern real R00248.kf;
  extern real R00248.kb;
  real R00256(time) katal;
  extern real R00256.kf;
  extern real R00256.kb;
  real R00575(time) katal;
  extern real R00575.kf;
  extern real R00575.kb;
  real R00497(time) katal;
  extern real R00497.kf;
  extern real R00497.kb;
  real R00768(time) katal;
  extern real R00768.kf;
  extern real R00768.kb;
  real R05578(time) katal;
  extern real R05578.kf;
  extern real R05578.kb;
  real R00355(time) katal;
  extern real R00355.kf;
  extern real R00355.kb;
  real R00253(time) katal;
  extern real R00253.kf;
  extern real R00253.kb;
  real R00094(time) katal;
  extern real R00094.kf;
  extern real R00094.kb;
  real R00713(time) katal;
  extern real R00713.kf;
  extern real R00713.kb;
  real R01231(time) katal;
  extern real R01231.kf;
  extern real R01231.kb;
  real R00243(time) katal;
  extern real R00243.kf;
  extern real R00243.kb;
  real R00114(time) katal;
  extern real R00114.kf;
  extern real R00114.kb;

  // equations
  when (time=time.min) Succinate = 0;
  Succinate:time = (R00713)/uVol;
  when (time=time.min) Succinate_space_semialdehyde = 0;
  Succinate_space_semialdehyde:time = (R01648 + -1*R00713)/uVol;
  when (time=time.min) space_2_minus_Oxoglutarate = 0;
  space_2_minus_Oxoglutarate:time = (R00248 + -1*R00355 + R00243 + R00114)/uVol;
  when (time=time.min) Glutathione_space_disulfide = 0;
  Glutathione_space_disulfide:time = (R00094)/uVol;
  when (time=time.min) Glutathione = 0;
  Glutathione:time = (R00497 + -1*R00094)/uVol;
  when (time=time.min) gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine = 0;
  gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine:time = (R00894 + -1*R00497)/uVol;
  when (time=time.min) L_minus_Glutamyl_minus_tRNA_space_Glu_space_ = 0;
  L_minus_Glutamyl_minus_tRNA_space_Glu_space_:time = (R05578)/uVol;
  when (time=time.min) space_4_minus_Aminobutanoate = 0;
  space_4_minus_Aminobutanoate:time = (R00261 + -1*R01648)/uVol;
  when (time=time.min) L_minus_Glutamine = 0;
  L_minus_Glutamine:time = (-1*R03652 + -1*R00257 + R01072 + -1*R00256 + -1*R00575 + -1*R00768 + R00253 + -1*R01231 + R00114)/uVol;
  when (time=time.min) L_minus_Glutamate = 0;
  L_minus_Glutamate:time = (-1*R00261 + -1*R00894 + -1*R00248 + R00256 + -1*R05578 + R00355 + -1*R00253 + -1*R00243 + -1*R00114)/uVol;
  when (time=time.min) space_5_minus_Phosphoribosylamine = 0;
  space_5_minus_Phosphoribosylamine:time = (-1*R01072)/uVol;
  when (time=time.min) NAD_plus_ = 0;
  NAD_plus_:time = (R00257)/uVol;
  when (time=time.min) GMP = 0;
  GMP:time = (R01231)/uVol;
  when (time=time.min) D_minus_Glucosamine_space_6_minus_phosphate = 0;
  D_minus_Glucosamine_space_6_minus_phosphate:time = (R00768)/uVol;
  when (time=time.min) Carbamoyl_space_phosphate = 0;
  Carbamoyl_space_phosphate:time = (R00575)/uVol;
  when (time=time.min) NH3 = 0;
  NH3:time = (R00256)/uVol;
  when (time=time.min) Glutaminyl_minus_tRNA = 0;
  Glutaminyl_minus_tRNA:time = (R03652)/uVol;
  R03652 = R03652.kf*L_minus_Glutamine - R03652.kb*Glutaminyl_minus_tRNA;
  R00257 = R00257.kf*L_minus_Glutamine - R00257.kb*NAD_plus_;
  R00261 = R00261.kf*L_minus_Glutamate - R00261.kb*space_4_minus_Aminobutanoate;
  R01648 = R01648.kf*space_4_minus_Aminobutanoate - R01648.kb*Succinate_space_semialdehyde;
  R00894 = R00894.kf*L_minus_Glutamate - R00894.kb*gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine;
  R01072 = R01072.kf*space_5_minus_Phosphoribosylamine - R01072.kb*L_minus_Glutamine;
  R00248 = R00248.kf*L_minus_Glutamate - R00248.kb*space_2_minus_Oxoglutarate;
  R00256 = R00256.kf*L_minus_Glutamine - R00256.kb*L_minus_Glutamate*NH3;
  R00575 = R00575.kf*L_minus_Glutamine - R00575.kb*Carbamoyl_space_phosphate;
  R00497 = R00497.kf*gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine - R00497.kb*Glutathione;
  R00768 = R00768.kf*L_minus_Glutamine - R00768.kb*D_minus_Glucosamine_space_6_minus_phosphate;
  R05578 = R05578.kf*L_minus_Glutamate - R05578.kb*L_minus_Glutamyl_minus_tRNA_space_Glu_space_;
  R00355 = R00355.kf*space_2_minus_Oxoglutarate - R00355.kb*L_minus_Glutamate;
  R00253 = R00253.kf*L_minus_Glutamate - R00253.kb*L_minus_Glutamine;
  R00094 = R00094.kf*Glutathione - R00094.kb*Glutathione_space_disulfide;
  R00713 = R00713.kf*Succinate_space_semialdehyde - R00713.kb*Succinate;
  R01231 = R01231.kf*L_minus_Glutamine - R01231.kb*GMP;
  R00243 = R00243.kf*L_minus_Glutamate - R00243.kb*space_2_minus_Oxoglutarate;
  R00114 = R00114.kf*L_minus_Glutamate - R00114.kb*L_minus_Glutamine*space_2_minus_Oxoglutarate;

  // variable properties
  default.sbmlRole="compartment";
  uVol.sbmlRole="compartment";
  E1_space_8_space_1_space_7.sbmlRole="species";
  E1_space_8_space_1_space_7.sbmlCompartment="uVol";
  E2_space_6_space_1_space_19.sbmlRole="species";
  E2_space_6_space_1_space_19.sbmlCompartment="uVol";
  E6_space_3_space_2_space_3.sbmlRole="species";
  E6_space_3_space_2_space_3.sbmlCompartment="uVol";
  E1_space_4_space_1_space_13.sbmlRole="species";
  E1_space_4_space_1_space_13.sbmlCompartment="uVol";
  E1_space_4_space_1_space_3.sbmlRole="species";
  E1_space_4_space_1_space_3.sbmlCompartment="uVol";
  E1_space_4_space_1_space_4.sbmlRole="species";
  E1_space_4_space_1_space_4.sbmlCompartment="uVol";
  E2_space_6_space_1_space_1.sbmlRole="species";
  E2_space_6_space_1_space_1.sbmlCompartment="uVol";
  E6_space_3_space_2_space_2.sbmlRole="species";
  E6_space_3_space_2_space_2.sbmlCompartment="uVol";
  E6_space_1_space_1_space_17.sbmlRole="species";
  E6_space_1_space_1_space_17.sbmlCompartment="uVol";
  E4_space_1_space_1_space_15.sbmlRole="species";
  E4_space_1_space_1_space_15.sbmlCompartment="uVol";
  E6_space_3_space_1_space_2.sbmlRole="species";
  E6_space_3_space_1_space_2.sbmlCompartment="uVol";
  E4_space_1_space_1_space_19.sbmlRole="species";
  E4_space_1_space_1_space_19.sbmlCompartment="uVol";
  E6_space_3_space_5_space_1.sbmlRole="species";
  E6_space_3_space_5_space_1.sbmlCompartment="uVol";
  E6_space_3_space_5_space_2.sbmlRole="species";
  E6_space_3_space_5_space_2.sbmlCompartment="uVol";
  E2_space_6_space_1_space_16.sbmlRole="species";
  E2_space_6_space_1_space_16.sbmlCompartment="uVol";
  E2_space_4_space_2_space_14.sbmlRole="species";
  E2_space_4_space_2_space_14.sbmlCompartment="uVol";
  E3_space_5_space_1_space_2.sbmlRole="species";
  E3_space_5_space_1_space_2.sbmlCompartment="uVol";
  E6_space_3_space_5_space_5.sbmlRole="species";
  E6_space_3_space_5_space_5.sbmlCompartment="uVol";
  E6_space_1_space_1_space_18.sbmlRole="species";
  E6_space_1_space_1_space_18.sbmlCompartment="uVol";
  E1_space_2_space_1_space_16.sbmlRole="species";
  E1_space_2_space_1_space_16.sbmlCompartment="uVol";
  Succinate.sbmlRole="species";
  Succinate.sbmlCompartment="uVol";
  Succinate_space_semialdehyde.sbmlRole="species";
  Succinate_space_semialdehyde.sbmlCompartment="uVol";
  space_2_minus_Oxoglutarate.sbmlRole="species";
  space_2_minus_Oxoglutarate.sbmlCompartment="uVol";
  Glutathione_space_disulfide.sbmlRole="species";
  Glutathione_space_disulfide.sbmlCompartment="uVol";
  Glutathione.sbmlRole="species";
  Glutathione.sbmlCompartment="uVol";
  gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine.sbmlRole="species";
  gamma_minus_L_minus_Glutamyl_minus_L_minus_cysteine.sbmlCompartment="uVol";
  L_minus_Glutamyl_minus_tRNA_space_Glu_space_.sbmlRole="species";
  L_minus_Glutamyl_minus_tRNA_space_Glu_space_.sbmlCompartment="uVol";
  space_4_minus_Aminobutanoate.sbmlRole="species";
  space_4_minus_Aminobutanoate.sbmlCompartment="uVol";
  L_minus_Glutamine.sbmlRole="species";
  L_minus_Glutamine.sbmlCompartment="uVol";
  L_minus_Glutamate.sbmlRole="species";
  L_minus_Glutamate.sbmlCompartment="uVol";
  space_5_minus_Phosphoribosylamine.sbmlRole="species";
  space_5_minus_Phosphoribosylamine.sbmlCompartment="uVol";
  NAD_plus_.sbmlRole="species";
  NAD_plus_.sbmlCompartment="uVol";
  GMP.sbmlRole="species";
  GMP.sbmlCompartment="uVol";
  D_minus_Glucosamine_space_6_minus_phosphate.sbmlRole="species";
  D_minus_Glucosamine_space_6_minus_phosphate.sbmlCompartment="uVol";
  Carbamoyl_space_phosphate.sbmlRole="species";
  Carbamoyl_space_phosphate.sbmlCompartment="uVol";
  NH3.sbmlRole="species";
  NH3.sbmlCompartment="uVol";
  Glutaminyl_minus_tRNA.sbmlRole="species";
  Glutaminyl_minus_tRNA.sbmlCompartment="uVol";
  R03652.sbmlRole="rate";
  R03652.kf.sbmlRole="rate";
  R03652.kb.sbmlRole="rate";
  R00257.sbmlRole="rate";
  R00257.kf.sbmlRole="rate";
  R00257.kb.sbmlRole="rate";
  R00261.sbmlRole="rate";
  R00261.kf.sbmlRole="rate";
  R00261.kb.sbmlRole="rate";
  R01648.sbmlRole="rate";
  R01648.kf.sbmlRole="rate";
  R01648.kb.sbmlRole="rate";
  R00894.sbmlRole="rate";
  R00894.kf.sbmlRole="rate";
  R00894.kb.sbmlRole="rate";
  R01072.sbmlRole="rate";
  R01072.kf.sbmlRole="rate";
  R01072.kb.sbmlRole="rate";
  R00248.sbmlRole="rate";
  R00248.kf.sbmlRole="rate";
  R00248.kb.sbmlRole="rate";
  R00256.sbmlRole="rate";
  R00256.kf.sbmlRole="rate";
  R00256.kb.sbmlRole="rate";
  R00575.sbmlRole="rate";
  R00575.kf.sbmlRole="rate";
  R00575.kb.sbmlRole="rate";
  R00497.sbmlRole="rate";
  R00497.kf.sbmlRole="rate";
  R00497.kb.sbmlRole="rate";
  R00768.sbmlRole="rate";
  R00768.kf.sbmlRole="rate";
  R00768.kb.sbmlRole="rate";
  R05578.sbmlRole="rate";
  R05578.kf.sbmlRole="rate";
  R05578.kb.sbmlRole="rate";
  R00355.sbmlRole="rate";
  R00355.kf.sbmlRole="rate";
  R00355.kb.sbmlRole="rate";
  R00253.sbmlRole="rate";
  R00253.kf.sbmlRole="rate";
  R00253.kb.sbmlRole="rate";
  R00094.sbmlRole="rate";
  R00094.kf.sbmlRole="rate";
  R00094.kb.sbmlRole="rate";
  R00713.sbmlRole="rate";
  R00713.kf.sbmlRole="rate";
  R00713.kb.sbmlRole="rate";
  R01231.sbmlRole="rate";
  R01231.kf.sbmlRole="rate";
  R01231.kb.sbmlRole="rate";
  R00243.sbmlRole="rate";
  R00243.kf.sbmlRole="rate";
  R00243.kb.sbmlRole="rate";
  R00114.sbmlRole="rate";
  R00114.kf.sbmlRole="rate";
  R00114.kb.sbmlRole="rate";
}